Members in Taiwan Pteridophyte Research Group are highlighted in GREEN

Chang, Z.-X.*, Kuo, L.-Y.*, Lu, P.-F., & Huang, Y.-M. 2020. New addition to the Asplenium normale complex (Aspleniaceae): an endemic forma in Taiwan. Taiwania, 65: 253–260. (* equally contibuted)

Kuo, L.-Y., Chang, Y.-H., Huang, Y.-H., Testo, W., Ebihara, A., Rouhan, G., Quintanillag, L. G., Watkins, J. E. J., Huang, Y.-M., & Li, F.-W. 2020. A global phylogeny of Stegnogramma ferns (Thelypteridaceae): generic and sectional revision, historical biogeography and evolution of leaf architecture. Cladistics, 26: 164–183.

Xu, K.-W., Zhang, L., Rothfels, C. J., Smith, A. R., Viane, R., Lorence, D., Wood, K. R., Chen, C.-W., Knapp, R., Zhou, L., Lu, N. T., Zhou, X.-M., Wei, H.-J., Fan, Q., Chen, S.-F., Cicuzzam, D., Gao, X.-F., Liao, W.-B., & Zhang, L.-B. 2020. A global plastid phylogeny of the fern genus Asplenium. Cladistics, 36: 22–71.

Lee, P. C., Chen, T. Y., Huagn, Y. M., Chu, Y., Chiou, W.L. 2020. Factors affecting longevity of spores of Pentarhizidium orientalei (Hook.) Hayata, an endangered fern in Taiwan. The International Journal of Plant Reproductive Biology 12 (2): 116–122. DOI 10.14787/ijprb.2020 12.2.


Chao, Y.-S. 2019. New record of Pteris esquirolii in Taiwan, with two similar new species, Pteris rugosifolia and Pteris subesquirolii. Syst. Bot., 44: 274–281.

Chao, Y.-S., Huang, Y.-F., Dong, S.-Y., Huang, Y.-M., & Liu, H.-Y. 2019. Bolbitis lianhuachihensis (Dryopteridaceae), a new species from Taiwan. PhytoKeys, 131: 69–81.

Chen, C.-W.*, Kuo, L.-Y.*, Huang, Y.-H., Hsu, T.-C., Dang, M. T., Luu, H. T., Li, C.-W., & Huang, Y.-M. 2019. A new species and a new record of Stegnogramma (Thelypteridaceae; Polypodiales) from southern Vietnam. Syst. Bot., 44: 768–774. (* equally contibuted)

Chen, C.-W., Lindsay, S., Yong, K. T., Mustapeng, A. M. A., Amoroso, V. B., Dang, V. D., & Huang, Y.-M. 2019. Clarification of two poorly known vittarioid ferns (Pteridaceae): Haplopteris angustissima and H. capillaris. Syst. Bot., 44: 483–493.

Ebihara, A., Nakato, N., Kuo, L.-Y., Miyazaki, H., & Serizawa, S. 2019. Allopolyploid origin and distribution range of Acystopteris taiwaniana (Cystopteridaceae: Polypodiales). Acta Phytotaxon. Geobot., 70: 19–28.

Hsu, T.-C., Huang, Y.-F., & Chao, Y.-S. 2019. Systematics of Hymenophyllum subgenus Mecodium (Hymenophyllaceae) in Taiwan. Syst. Bot., 44: 753–767.

Huang, Y.-M., Chang, Y.-H., & Chiou, W.-L. 2019. Edible ferns and lycophytes in Asia. Fern Gaz., 21: 45–68.

Huang, Y.-M., Chen, C.-W., Chiou, W.-L., Chang, H.-M., & Lin, H.-Y. 2019. Ex situ conservation of threatened ferns and lycophytes in Taiwan, aspect of reproductive biology. Int. J. Plant Reprod. Biol., 11: 121–127.

Huang, Y.-M., Lee, P.-S., & Chiou, W.-L. 2019. Phenology and leaf lifespan of an endangered Angiopteris somae (Marattiaceae). Indian Fern J., 36: 133–153.

Kirschner, R., Lee, P.-H., & Huang, Y.-M. 2019. Diversity of fungi on Taiwanese fern plants : review and new discoveries. Taiwania, 64: 163–175.

TPG: Kuo, L.-Y.*, Hsu, T.-C.*, Chao, Y.-S., Liou, W.-T., Chang, H.-M., Chen, C.-W., Huang, Y.-M., Li, F.-W., Huang, Y.-F., Shao, W., Lu, P.-F., Chen, C.-W., Chang, Y.-H., & Chiou, W.-L. 2019. Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania, 64: 367–395. (* equally contibuted)

Kuo, L.-Y., & Li, F.-W. 2019. A roadmap for fern genome sequencing. Am. Fern J., 109: 212–223.

Xu, K., Chen, C.-W., Kamau, P., Liao, W.-B., & Zhang, L.-B. 2019. Four new species of the fern genus Hymenasplenium (Aspleniaceae) from Africa and Asia. Phytotaxa, 416: 34–42.


Chao, Y.-S., & Huang, Y.-M. 2018. Spore morphology and its systematic implication in Pteris (Pteridaceae). PLoS ONE, 13: e0207712.

Chen, C.-W., Lindsay, S., Mynssen, C. M., Kuo, L.-Y., Amoroso, V. B., & Huang, Y.-M. 2018. A new Haplopteris species from the Philippines and clarification of the status of H. amboinensis in China and Indochina. Gard. Bull. Singapore, 70: 177–190.

Chen, C.-W.*, Rothfels, C. J.*, Mustapeng, A. M. A., Gubilil, M., Karger, D. N., Kessler, M., & Huang, Y.-M. 2018. End of an enigma: Aenigmopteris belongs in Tectaria (Tectariaceae: Polypodiopsida). J. Plant Res., 131: 67–76. (* equally contibuted)

Huang, Y.-M., Yap, Y.-Q., & Li, C.-W. 2018. A semiaquatic but desiccation-tolerant plant, Isoetes taiwanensis DeVol (Isoetaceae; Lycophyta). Int. J. Plant Reprod. Biol., 10: 10–13.

Kuo, L.-Y., Ebihara, A., Hsu, T.-C., Rouhan, G., Huang, Y.-M., Wang, C.-N., Chiou, W.-L., & Kato, M. 2018. Infrageneric revision of the fern genus Deparia (Athyriaceae, Aspleniineae, Polypodiales). Syst. Bot., 43: 645–655.

Kuo, L.-Y., Ebihara, A., Kato, M., Rouhan, G., Ranker, T. A., Wang, C.-N., & Chiou, W.-L. 2018. Morphological characterization of infra-generic lineages in Deparia (Athyriaceae: Polypodiales). Cladistics, 34: 78–92.

Kuo, L.-Y., Qi, X., Ma, H., & Li, F.-W. 2018. Order-level fern plastome phylogenomics: new insights from Hymenophyllales. Am. J. Bot., 105: 1545–1555.

Kuo, L.-Y., Tang, T.-Y., Chiou, W.-L., Li, F.-W., Huang, Y.-M., & Wang, C.-N. 2018. Organelle genome inheritances in Deparia ferns (Athyriaceae, Aspleniineae, Polypodiales). Front. Plant Sci., 9: Article 486.

Lee, P.-H., Huang, Y.-M., & Chiou, W.-L. 2018. Fern Phenology. Pp. 381–399 in: H. Fernández (ed.), Current Advances in Fern Research. Cham: Springer International Publishing.

Li, F.-W., Brouwer, P., Carretero-Paulet, L., Cheng, S., Vries,, Delaux, P.-M., Eily, A., Koppers, N., Kuo, L.-Y., Li, Z., Simenc, M., Small, I., Wafula, E., Angarita, S., Barker, M. S., Braeutigam, A., DePamphilis, C., Gould, S., Hosmani, P. S., Huang, Y.-M., Huettel, B., Kato, Y., Liu, X., Maere, S., McDowell, R., Mueller, L. A., Rensing, S. A., Robison, T., Rothfels, C. J., Sigel, E. M., Song, Y., Timilsina, P. R., Peer, Y.Van de, Wang, H., Wilhelmsson, P. K. I., Wolf, P. G., Xu, X., Der, J. P., Schluepmann, H., Wong, G. K.-S., & Pryer, K. M. 2018. Fern genomes elucidate land plant evolution and cyanobacterial symbioses. Nat. Plants, 4: 460–472.

Qi, X., Kuo, L.-Y., Guo, C., Li, H., Li, Z., Qi, J., Wang, L., Hu, Y., Xiang, J., Zhang, C., Guo, J., Huang, C.-H., & Ma, H. 2018. A well-resolved fern nuclear phylogeny reveals the evolution history of numerous transcription factor families. Mol. Phylogenet. Evol., 127: 961–977.

Song, M.*, Kuo, L.-Y.*, Huiet, L., Pryer, K. M., Rothfels, C. J., & Li, F.-W. 2018. A novel chloroplast gene reported for flagellate plants. Am. J. Bot., 105: 117–121. (* equally contibuted)


Chao, Y.-S., Ebihara, A., Chiou, W.-L., & Huang, Y.-M. 2017. Pteris latipinna sp. nov. (Pteridaceae), a new species segregated from Pteris fauriei. PhytoKeys, 85: 95–108.

Chao, Y.-S., Mustapeng, A. M. A., Chen, C.-W., & Chiou, W.-L. 2017. Pteris borneensis (Pteridaceae), a new Species from Borneo, with re-circumscription of Pteris decrescens and Pteris parviloba. Syst. Bot., 42: 724–732.

Chen, C.-W., Lindsay, S., Kuo, L.-Y., Fraser-Jenkins, C. R., Ebihara, A., Luu, H. T., Park, C. W., Chao, Y.-S., Huang, Y.-M., & Chiou, W.-L. 2017. A systematic study of East Asian vittarioid ferns (Pteridaceae, Vittarioideae). Bot. J. Linn. Soc., 183: 545–560.

Chen, C.-W., Perrie, L., Glenny, D., & Chiou, W.-L. 2017. Sol amazing: lycophytes and ferns of the Solomon Islands. Taichung, Taiwan: National Museum of Natural Science.

Chen, C.-W.*, Sundue, M.*, Kuo, L.-Y., Teng, W.-C., & Huang, Y.-M. 2017. Phylogenetic analyses place the monotypic Dryopolystichum within Lomariopsidaceae. PhytoKeys, 78: 83–107. (* equally contibuted)

Dong, S.-Y., Chen, C.-W., Tan, S.-S., Zhao, H.-G., Zuo, Z.-Y., Chao, Y.-S., & Chang, Y.-H. 2017. New insights on the phylogeny of Tectaria (Tectariaceae), with special reference to Polydictyum as a distinct lineage. J. Syst. Evol., 56: 139–147.

Fujiwara, T., Uehara, A., Iwashina, T., Matsumoto, S., Chang, Y.-H., Chao, Y.-S., & Watano, Y. 2017. Allotetraploid cryptic species in Asplenium normale in the Japanese Archipelago, detected by chemotaxonomic and multi-locus genotype approaches. Am. J. Bot., 104: 1390–1406.

Hori, K., Kuo, L.-Y., Chiou, W.-L., Ebihara, A., & Murakami, N. 2017. Geographical distribution of sexual and apogamous types of Dryopteris formosana and Dryopteris varia (Dryopteridaceae) in Taiwan. Acta Phytotaxon. Geobot., 68: 23–32.

Kuo, L.-Y.*, Chen, C.-W.*, Shinohara, W., Ebihara, A., Kudoh, H., Sato, H., Huang, Y.-M., & Chiou, W.-L. 2017. Not only in the temperate zone: independent gametophytes of two vittarioid ferns (Pteridaceae, Polypodiales) in East Asian subtropics. J. Plant Res., 130: 255–262. (* equally contibuted)

Kuo, L.-Y., Huang, Y.-J., Chang, J., Chiou, W.-L., & Huang, Y.-M. 2017. Evaluating the spore genome sizes of ferns and lycophytes: a flow cytometry approach. New Phytol., 213: 1974–1983.

Kuo, L.-Y., & Huang, Y.-M. 2017. Determining genome size from spores of seedless vascular plants. Bio-Protocol, 7: e2322.

Pinson, J. B., Chambers, S. M., Nitta, J. H., Kuo, L.-Y., & Sessa, E. B. 2017. The separation of generations: biology and biogeography of long-lived sporophyteless fern gametophytes. Int. J. Plant Sci., 178: 1–18.

Tseng, M.-H., Lin, K.-H., Huang, Y.-J., Chang, Y.-L., Huang, S.-C., Kuo, L.-Y., & Huang, Y.-M. 2017. Detection of chlorophylls in spores of seven ferns. J. Plant Res., 130: 407–416.

You, C.*, Cui, J.*, Wang, H., Qi, X., Kuo, L.-Y., Ma, H., Gao, L., Mo, B., & Chen, X. 2017. Conservation and divergence of small RNA pathways and microRNAs in land plants. Genome Biol., 18: 158. (* equally contibuted)

Zhou, X.-M., Chen, C.-W., & Zhang, L.-B. 2017. Pyrrosia annamensis comb. nov. (Polypodiaceae) from southeast asia and lectotypification of Cyclophorus rhomboidalis. Phytotaxa, 309: 90–94.

Zhou, X.-M., Zhang, L., Chen, C.-W., Li, C.-X., Huang, Y.-M., Chen, D.-K., Lu, N. T., Cicuzza, D., Knapp, R., Luong, T. T., Nitta, J. H., Gao, X.-F., & Zhang, L.-B. 2017. A plastid phylogeny and character evolution of the Old World fern genus Pyrrosia (Polypodiaceae) with the description of a new genus: Hovenkampia (Polypodiaceae). Mol. Phylogenet. Evol., 114: 271–294.


Amoroso, V. B., Chen, C.-W., Coritico, F. P., Lu, P.-F., Alcala, E. L., & Chiou, W.-L. 2016. Guide to lycophytes and ferns of Balinsasayao, Negros, Philippines. Pingtung, Taiwan: KBCC press.

Chen, C.-W., Schuettpelz, E., Lindsay, S., & Middleton, D. J. 2016. Proposal to conserve the name Haplopteris against Monogramma (Pteridaceae). Taxon, 65: 884–885.

Chiou, W.-L., Hu, J.-M., Chen, C.-H., Chien, C.-F., Huang, H.-C., & Yang, T.-Y. A. 2016. Field Guide to the Plants of Solomon Islands. Taichung, Taiwan: National Museum of Natural Science.

Dong, S.-Y., Zuo, Z.-Y., Chao, Y.-S., Damas, K., & Sule, B. 2016. New species of the fern genus Lindsaea (Lindsaeaceae) from New Guinea with notes on the phylogeny of L. sect. Synaphlebium. PLoS ONE, 11: e0163686.

Ebihara, A., Nakato, N., Amoroso, V. B., Hidayat, A., & Kuo, L.-Y. 2016. The Japanese endemic Monachosorum arakii Tagawa (Dennstaedtiaceae) is a relict 'inter-national' hybrid: a reassessment of the Monachosorum species. Syst. Bot., 41: 586–595.

Kuo, L.-Y., Chang, Y.-H., Glowienka, J. M. O., Amoroso, V. B., Dong, S.-Y., Kao, T.-T., Wang, C.-N., & Chiou, W.-L. 2016. A revised framework of Dryopteris subg. Nothoperanema (Dryopteridaceae) inferred from phylogenetic evidence, with descriptions of two new sections. Syst. Bot., 41: 596–605.

Kuo, L.-Y., Ebihara, A., Shinohara, W., Rouhan, G., Wood, K. R., Wang, C.-N., & Chiou, W.-L. 2016. Historical biogeography of the fern genus Deparia (Athyriaceae) and its relation with polyploidy. Mol. Phylogenet. Evol., 104: 123–134.

Kuo, L.-Y., Hsu, T.-C., Chang, Y.-H., Huang, Y.-M., & Chiou, W.-L. 2016. Systematics of the Adiantum philippense complex (Pteridaceae, Polypodiales) in Taiwan. Phytotaxa, 263: 113–121.

Lee, P.-H., Chen, S.-W., Chiou, W.-L., Huang, Y.-M., & Liu, H.-Y. 2016. Phenology of 13 fern species in a tropical monsoon forest of southern Taiwan. Int. J. Plant Reprod. Biol., 8: 87–97.

Li, F.-W.*, Kuo, L.-Y.*, Chang, Y.-H., Hsu, T.-C., Hung, H.-C., Chiou, W.-L., Rothfels, C. J., & Huang, Y.-M. 2016. Asplenium pifongiae (Aspleniaceae: Polypodiales), a new species from Taiwan. Syst. Bot., 41: 24–31. (* equally contibuted)

Li, F.-W., Kuo, L.-Y., Pryer, K. M., & Rothfels, C. J. 2016. Genes translocated into the plastid inverted repeat show decelerated substitution rates and elevated GC content. Genome Biol. Evol., 8: 2452–2458.

Liou, W.-T., Lu, P.-F., Huang, Y.-M., & Yang, C.-K. 2016. The confirmation of three Dryopteris species (Dryopteridaceae) in Taiwan. J. Exp. For. Natl. Taiwan Univ., 30: 293–302.

PPG I. 2016. A community-derived classification for extant lycophytes and ferns. J. Syst. Evol., 54: 563–603.

Tsutsumi, C., Chen, C.-W., Larsson, A., Hirayama, Y., & Kato, M. 2016. Phylogeny and classification of Davalliaceae on the basis of chloroplast and nuclear markers. Taxon, 65: 1236–1248.


Chang, Y.-H., Chang, H.-M., Hung, H.-C., Teng, W.-C., Chen, C.-W., & Hsu, T.-C. 2015. Rediscovery of Vaginularia trichoidea Fée (Pteridaceae) in Taiwan. Taiwan J. Biodivers., 17: 59–65.

Chang, Y.-L., Tseng, M.-H., Chiou, W.-L., & Huang, Y.-M. 2015. Effect of temperature on the viability, lifespan and vigour, of chlorophyllous spores of Osmundastrum cimmamomeum (Osmundaceae). Fern Gaz., 20: 55–64.

Chao, Y.-S., Liu, H.-Y., & Chiou, W.-L. 2015. Taxonomic revision of the Pteris cadieri complex (Pteridaceae). Phytotaxa, 230: 130–150.

Chen, C.-W., Nitta, J. H., Fanerii, M., Yang, T.-Y. A., Pitisopa, F., Li, C.-W., & Chiou, W.-L. 2015. Antrophyum solomonense (Pteridaceae), a new Species from the Solomon Islands, and its systematic position based on phylogenetic analysis. Syst. Bot., 40: 645–651.

Chiu, T.-Y., Wang, H.-H., Kuo, Y.-L., Tomonori, K., Chiou, W.-L., & Huang, Y.-M. 2015. Ecophysiological characteristics of three Cyathea species in Northeastern Taiwan. Taiwan J. For. Sci., 30: 147–155.

Hsu, T.-C., Hung, H.-C., Lin, C.-Y., Chiou, W.-L., Huang, Y.-M., & Chang, Y.-H. 2015. New additions to the fern flora of Taiwan (3). Taiwan J. For. Sci., 30: 131–137.

Huang, Y.-M., Amoroso, V. B., Coritico, F. P., Ko, C.-W., Kao, T.-T., Callado, J. R., & Chiou, W.-L. 2015. Reproductive biology of Aglaomorpha cornucopia (Copel.) M. C. Roos, a rare and endemic fern from the Philippines. Am. Fern J., 105: 31–44.

Chang, Y.-H., Chiou, W.-L., Huang, Y.-M., Shen, B.-N., Lin, C.-Y., & Hsu, T.-C. 2014. New additions to the fern flora of Taiwan (2). Taiwan J. Biodivers., 16: 263–272.

Chao, Y.-S., Rouhan, G., Amoroso, V. B., & Chiou, W.-L. 2014. Molecular phylogeny and biogeography of the fern genus Pteris (Pteridaceae). Ann. Bot., 114: 109–124.

Chen, C.-W., Ebihara, A., Chiou, W.-L., & Li, C.-W. 2014. Haplopteris yakushimensis (Pteridaceae, Vittarioideae), a new species from Yakushima Island, Japan. Phytotaxa, 156: 229–234.

Chen, C.-W., Ngan, L. T., Hidayat, A., Evangelista, L., Nooteboom, H. P., & Chiou, W.-L. 2014. First insights into the evolutionary history of the Davallia repens complex. Blumea, 59: 49–58.

Ding, H.-H.*, Chao, Y.-S.*, Callado, J. R., & Dong, S.-Y. 2014. Phylogeny and character evolution of the fern genus Tectaria (Tectariaceae) in the Old World inferred from chloroplast DNA sequences. Mol. Phylogenet. Evol., 80: 66–78. (* equally contibuted)

Ebihara, A., Nakato, N., Matsumoto, S., Chao, Y.-S., & Kuo, L.-Y. 2014. Cytotaxonomic studies on thirteen ferns of Taiwan. Bull. Natl. Museum Nat. Sci. Ser. B, 40: 19–28

Hsu, T.-C., Lin, C.-Y., Chen, C.-W., Chiou, W.-L., & Chang, Y.-H. 2014. Confirmation of the occurrence and distribution of three fern species in Taiwan. Taiwan J. For. Sci., 29: 85–95.

Kao, T.-T., Chiou, W.-L., Hsu, S.-Y., Chen, C.-M., Chao, Y.-S., & Huang, Y.-M. 2014. Hybrid origin of Nephrolepis × hippocrepicis Miyam. (Nephrolepidaceae). Int. J. Plant Reprod. Biol., 6: 1–14.

Lindsay, S., & Chen, C.-W. 2014. Three new combinations in Haplopteris (Pteridaceae subfam. Vittarioideae). Gard. Bull. Singapore, 66: 169–171.

Sundue, M. A., Parris, B. S., Ranker, T. A., Smith, A. R., Fujimoto, E. L., Zamora-Crosby, D., Morden, C. W., Chiou, W.-L., Chen, C.-W., Rouhan, G., Hirai, R. Y., & Prado, J. 2014. Global phylogeny and biogeography of grammitid ferns (Polypodiaceae). Mol. Phylogenet. Evol., 81: 195–206.

Zhang, W.-Y.*, Kuo, L.-Y.*, Li, F.-W., Wang, C.-N., & Chiou, W.-L. 2014. The hybrid origin of Adiantum meishanianum (Pteridaceae): a rare and endemic species in Taiwan. Syst. Bot., 39: 1034–1041. (* equally contibuted)

Chang, Y.-H., Kuo, L.-Y., Chiou, W.-L., Chen, C.-W., Chang, H.-M., & Amoroso, V. B. 2013. Stenolepia Alderw. (Dryopteridaceae), a fern genus new to the Philippines. Philipp. J. Sci., 142: 83–88.

Chang, Y.-L., Hsieh, C.-L., Huang, Y.-M., Chiou, W.-L., Kuo, Y.-H., & Tseng, M.-H. 2013. Modified method for determination of sulfur metabolites in plant tissues by stable isotope dilution-based liquid chromatography-electrospray ionization-tandem mass spectrometry. Anal. Biochem., 442: 24–33.

Chao, Y.-S., Ebihara, A., Chang, Y.-H., Jiang, R.-H., Ngan, L. T., & Chiou, W.-L. 2013. New distribution of Pteris kawabatae Sa. Kurata and re-circumscription of Pteris arisanensis Tagawa. Taiwania, 58: 305–310.

Chen, C.-W., Huang, Y.-M., Kuo, L.-Y., Chang, Y.-H., Liu, Y.-C., & Chiou, W.-L. 2013. A new vittarioid fern species, Haplopteris heterophylla (Pteridaceae). Syst. Bot., 38: 901–909.

Chen, C.-W., Huang, Y.-M., Kuo, L.-Y., Nguyen, Q. D., Luu, H. T., Callado, J. R., Farrar, D. R., & Chiou, W.-L. 2013. TrnL-F is a powerful marker for DNA identification of field vittarioid gametophytes (Pteridaceae). Ann. Bot., 111: 663–673.

Chiu, T.-Y., Chiou, W.-L., & Huang, Y.-M. 2013. Phenological differences between sterile and fertile fronds ina dimorphic fern, Osmundastrum cinnamomeum (L.) C.Presl (Osmundaceae). Taiwan J. Biodivers., 15: 311–322.

Ding, H.-H., Chao, Y.-S., & Dong, S.-Y. 2013. Taxonomic novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa, 122: 61–64.

Fu, C.-H., Hsieh, H.-M., Chen, C.-Y., Chang, T.-T., Huang, Y.-M., & Ju, Y.-M. 2013. Ophiodiaporthe cyatheae gen. et sp. nov., a diaporthalean pathogen causing a devastating wilt disease of Cyathea lepifera in Taiwan. Mycologia, 105: 861–872.

Liou, W.-T., Yang, C.-K., Wang, Y.-N., Chang, P.-C., & Huang, Y.-M. 2013. The investigation of lycophytes and monilophytes in Xitou Tract, Experimental Forest, National Taiwan University. J. Exp. For. Natl. Taiwan Univ., 27: 33–48.

Moore, S.-J., Parris, B. S., Kao, T.-T., Lu, P.-F., & Chiou, W.-L. 2013. Xiphopterella devolii (Polypodiaceae), a new species and newly recorded genus in Taiwan. Bot. Stud., 54: 24.

Chang, H.-M., Chiou, W.-L., & Wang, J.-C. 2012. Fern Flora of Taiwan, Selaginellaceae. Nantou, Taiwan: Endemic Species Research Institute.

Chang, Y.-H., Chiou, W.-L., Lu, P.-F., & Hsu, T.-C. 2012. A rediscovery of Microlepia platyphylla (Don) J. Sm. (Dennstaedtiaceae) from Taiwan. Taiwan J. Biodivers., 14: 15–21.

Chang, Y.-H., Wu, W.-H., Chen, C.-W., Lu, P.-F., & Huang, Y.-M. 2012. Two taxa of ferns newly recorded to Taiwan. Taiwan J. Biodivers., 14: 81–90.

Chao, Y.-S., Chang, Y.-H., Ngan, L.-T., Truong, D.Van, Liu, H.-Y., & Chiou, W.-L. 2012. Different ploids of Pteris grevilleana Wall. ex Agardh var. ornata Alderw. (Pteridaceae) in Taiwan and Vietnam. Taiwania, 57: 278–282.

Chao, Y.-S., Dong, S.-Y., Chiang, Y.-C., Liu, H.-Y., & Chiou, W.-L. 2012. Extreme multiple reticulate origins of the Pteris cadieri complex (Pteridaceae). Int. J. Mol. Sci., 13: 4523–4544.

Chao, Y.-S., Liu, H.-Y., Chiang, Y.-C., & Chiou, W.-L. 2012. Polyploidy and speciation in Pteris (Pteridaceae). J. Bot., 2012: 1–7.

Chen, C.-W., Kuo, L.-Y., Wang, C.-N., & Chiou, W.-L. 2012. Development of PCR primer sets for intron 1 of the low-copy gene LEAFY in Davalliaceae. Am. J. Bot., 99: e223-225.

Dong, S.-Y., Wei, L.-L., & Chao, Y.-S. 2012. A new species of Asplenium section Thamnopteris (Aspleniaceae) from Indonesia. Blumea, 57: 190–194.

Ebihara, A., Fraser-Jenkins, C. R., Parris, B. S., Zhang, X.-C., Yang, Y.-H., Chiou, W.-L., Chang, H.-M., Lindsay, S., Middleton, D., Kato, M., Praptosuwiryo, T. N., Amoroso, V. B., Barcelona, J. F., Ranil, R. H. G., Park, C.-H., Murakami, N., & Hoya, A. 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bull. Natl. Museum Nat. Sci. Ser. B, Bot., 38: 93–119.

Ebihara, A., & Kuo, L.-Y. 2012. East and Southeast Asian pteridophyte flora and DNA barcoding. Pp. 321–327 in: S.Nakano, T.Yahara, & T.Nakashizuka (eds.), The Biodiversity Observation Network in the Asia-Pacific Region: Toward Further Development of Monitoring, Ecological Research Monographs. Tokyo: Springer Japan.

Rothfels, C. J.*, Larsson, A.*, Kuo, L.-Y.*, Korall, P., Chiou, W.-L., & Pryer, K. M. 2012. Overcoming deep roots, fast rates, and short internodes to resolve the ancient rapid radiation of eupolypod II ferns. Syst. Biol., 61: 490–509. (* equally contibuted)

Rothfels, C. J.*, Sundue, M. A.*, Kuo, L.-Y., Larsson, A., Kato, M., Schuettpelz, E., & Pryer, K. M. 2012. A revised family-level classification for eupolypod II ferns (Polypodiidae: Polypodiales). Taxon, 61: 515–533. (* equally contibuted)

Chang, H.-M., Lu, P.-F., Hsu, T.-C., & Chiou, W.-L. 2011. Selaginella devolii (Selaginellaceae), a new species from Taiwan. Blumea, 56: 21–23.

Huang, Y.-M., Hsu, S., Hsieh, T.-H., Chou, H.-M., & Chiou, W.-L. 2011. Three Pteris species (Pteridaceae: Pteridophyta) reproduce by apogamy. Bot. Stud., 52: 79–87.

Huang, Y.-M., Huang, M.-H., Chen, C.-M., & Chiou, W.-L. 2011. Stipule propagation in five marattioid species native to Taiwan (Marattiaceae; Pteridophyte). Pp. 127–134 in: Working with Ferns. New York: Springer New York.

Huang, Y.-M., Liu, Y.-C., Lu, P.-F., & Chiou, W.-L. 2011. Gametophytes and young sporophytes of Pentarhizidium orientalis (Hook.) Hayata (Onocleaceae), a rare fern in Taiwan. Int. J. Plant Reprod. Biol., 3: 7–14.

Kuo, L.-Y.*, Li, F.-W.*, Chiou, W.-L., & Wang, C.-N. 2011. First insights into fern matK phylogeny. Mol. Phylogenet. Evol., 59: 556–566.

Li, F.-W., Kuo, L.-Y., Rothfels, C. J., Ebihara, A., Chiou, W.-L., Windham, M. D., & Pryer, K. M. 2011. RbcL and matK earn two thumbs up as the core DNA barcode for ferns. PLoS ONE, 6: e26597.

Liu, Y.-C., Chiou, W.-L., & Kato, M. 2011. Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae). Taxon, 60: 824–830.

Wu, W.-H., Chang, Y.-H., Chiou, W.-L., & Huang, Y.-M. 2011. Ferns and lycophytes in southwestern Yangmingshan National Park. Natl. Park Bull., 21: 35–46.


Chao, Y.-S., Liu, H.-Y., Huang, Y.-M., & Chiou, W.-L. 2010. Reproductive traits of Pteris cadieri and P. grevilleana in Taiwan: implications for their hybrid origin. Bot. Stud., 51: 209–216.

Kao, Y.-C., Chiou, T.-Y., Chu, M.-K., Chiou, W.-L., & Huang, Y.-M. 2010. Study of the fertility in Osmunda cinnamomea L. at Cao-Pi area. J. Natl. Taiwan Museum, 63: 15–25.

Li, F.-W.*, Kuo, L.-Y.*, Huang, Y.-M., Chiou, W.-L., & Wang, C.-N. 2010. Tissue-direct PCR, a rapid and extraction-free method for barcoding of ferns. Mol. Ecol. Resour., 10: 92–95. (* equally contibuted)

Chang, H.-M., Chen, J.-F., Lu, P.-F., Moore, S.-J., Wang, J.-C., & Chiou, W.-L. 2009. Supplements to the lycophytes in Taiwan (I): a newly recorded species Selaginella lutchuensis Koidz. (Selaginellaceae). Taiwania, 54: 88–92.

Chang, H.-M., Chiou, W.-L., & Wang, J.-C. 2009. Molecular evidence for genetic heterogeneity and the hybrid origin of Acrorumohra subreflexipinna from Taiwan. Am. Fern J., 99: 61–77.

Huang, Y.-M., Hsu, S.-Y., Huang, M.-H., & Chiou, W.-L. 2009. Reproductive biology of three cheilanthoid ferns in Taiwan. Int. J. Plant Reprod. Biol., 1: 109–116.

Lee, P.-H., Chiou, W.-L., & Huang, Y.-M. 2009. Phenology of three Cyathea (Cyatheaceae) ferns in northern Taiwan. Taiwan J. For. Sci., 24: 233–242.

Lee, P.-H., Lin, T.-T., & Chiou, W.-L. 2009. Phenology of 16 species of ferns in a subtropical forest of northeastern Taiwan. J. Plant Res., 122: 61–67.

Liu, Y.-C., Chiou, W.-L., & Liu, H.-Y. 2009. Fern Flora of Taiwan: Athyrium. Taiwan For. Res. Inst. Ext., 198: 1–112.

Liu, Y.-C., Huang, Y.-M., & Chiou, W.-L. 2009. Validation of the name Adiantum meishanianum (Pteridaceae), a species endemic to Taiwan. Novon, 19: 59–61.

Cheng, Y.-P., Liu, S.-H., Huang, Y.-M., Chen, C.-W., & Chiou, W.-L. 2008. Allozyme variations of a widespread tree fern, Alsophila spinulosa (Hook.) Tryon (Cyatheaceae), in Taiwan. Taiwan J. For. Sci., 23: 21–34.

Hsieh, T.-H., Chang, Y.-H., Chiou, W.-L., & Huang, Y.-M. 2008. Chromosome numbers of five species of the Marattiaceae in Taiwan. Taiwan J. Biodivers., 23: 335–339.

Huang, Y.-M., Chou, H.-M., Liu, C.-J., & Chiou, W.-L. 2008. The correlation of plant size and fertility of Asplenium nidus L. (Aspleniaceae; Pteridophyta). Bio Formosa, 43: 77–83.

Huang, Y.-W., Chiang, T.-Y., & Chiou, W.-L. 2008. Isolation and characterization of microsatellite loci of Lycopodium fordii Bak. (Lycopodiaceae, Pteridophyta). Conserv. Genet., 9: 775–777.

Kao, T.-T., Chen, S.-J., Chiou, W.-L., Chuang, Y.-C., & Kuo-Huang, L.-L. 2008. Various microscopic methods for investigating the venuloid idioblasts of Pteris grevilleana Wall. Taiwania, 53: 394–399.

Lee, P.-H., Huang, Y.-M., & Chiou, W.-L. 2008. The phenology of Osmunda claytoniana L. in the Tataka area, central Taiwan. Taiwan J. For. Sci., 23: 85–93.

Liu, Y.-C., Fraser-Jenkins, C., Amoroso, V. B., & Chiou, W.-L. 2008. Athyrium erythropodum (Woodsiaceae, pteridophyta), a new Philippine record. Blumea, 447–451.

Chang, H.-M., Chiang, H.-L., Lu, P.-F., Wang, J.-C., & Chiou, W.-L. 2007. Supplements to the pteridophyte in Taiwan (III): a newly recorded species Dryopteris championii (Benth.) C. Chr. ex Ching (Dryopteridaceae). Taiwania, 52: 238–242.

Chou, H.-M., Huang, Y.-M., Wong, S.-L., Hsieh, T.-H., Hsu, S.-Y., & Chiou, W.-L. 2007. Observations on gametophytes and juvenile sporophytes of Archangiopteris somai Hayata (Marattiaceae), an endangered fern in Taiwan. Bot. Stud., 48: 205–213.

Huang, Y.-M., Chou, H.-M., Wang, J.-C., & Chiou, W.-L. 2007. The distribution and habitats of the Pteris fauriei complex in Taiwan. Taiwania, 52: 49–58.

Moore, S.-J., Wang, J.-C., & Chiou, W.-L. 2007. Pteridophytes. in: C.-I.Peng, D. E.Boufford, T.Takaso, &T.-Y.Chiang (eds.), A Selection of Plants from Iriomote Island, Japan. Nantou, Taiwan: Endemic Species Research Institute; Taipei, Taiwan: Research Center for Biodiversity, Academic Sinica.

Chang, H.-M., Moore, S.-J., Chiou, W.-L., & Wang, J.-C. 2006. Supplements to the pteridophytes in Taiwan (II): a newly recorded species Diplazium crassiusculum Ching (Athyriaceae). Taiwania, 51: 287–292.

Chiou, W.-L., Huang, Y.-M., & Chen, C.-M. 2006. Conservation of two endangered ferns, Archangiopteris somai and A. itoi (Marattiaceae: Pteridophyta), by propagation from stipules. Fern Gaz., 17: 271–278.

Chiou, W.-L., Huang, Y.-M., Hsieh, T.-H., & Hsu, S.-Y. 2006. Diplazium megaphyllum (Bak.) Christ, a rare fern in Taiwan, reproduces by apogamy. Taiwan J. For. Sci., 21: 39–47.

Huang, Y.-M., Chou, H.-M., Hsieh, T.-H., Wang, J.-C., & Chiou, W.-L. 2006. Cryptic characteristics distinguish diploid and triploid varieties of Pteris fauriei (Pteridaceae). Can. J. Bot., 84: 261–268.


Chiou, W.-L., Martin, C., Lin, T.-C., & Hsu, C.-C. 2005. Ecophysiological differences between sterile and fertile fronds of the subtropical epiphytic fern Pyrrosia lingua (Polypodiaceae) in Taiwan. Am. Fern J., 95: 131–140.

Chiou, W.-L., & Moore, S.-J. 2005. A history of taxonomic studies of the vascular plants of Taiwan. J. Japanese Bot., 80: 309–312.

Huang, Y.-M., Chou, H.-M., & Chiou, W.-L. 2004. Density affects gametophyte growth and sexual expression of Osmunda cinnamomea (Osmundaceae: Pteridophyta). Ann. Bot., 94: 229–232.

Ko, J.-W., Huang, Y.-M., & Chiou, W.-L. 2004. Reproductive biology of Aglaomorpha meyeniana (Polypodiaceae; Pteridophyta). Bio Formosa, 39: 83–91.

Martin, C. E., Lin, T.-C., Hsu, C.-C., Lin, S.-H., Lin, K.-C., Hsia, Y.-J., & Chiou, W.-L. 2004. Ecophysiology and plant size in a tropical epiphytic fern, Asplenium nidus, in Taiwan. Int. J. Plant Sci., 165: 65–72.

Martin, C. E., Lin, T.-C., Lin, K.-C., Hsu, C.-C., & Chiou, W.-L. 2004. Causes and consequences of high osmotic potentials in epiphytic higher plants. J. Plant Physiol., 161: 1119–1124.

Chang, H.-M., Chiou, W.-L., & Wang, J.-C. 2003. Supplements to the pteridophytes in Taiwan (I): Dryopteris decipiens (Hook.) Kuntze (Dryopteridaceae). Taiwania, 48: 197–202.

Chiang, T.-Y., Chiang, Y.-C., Chou, C.-H., Cheng, Y.-P., & Chiou, W.-L. 2002. Phylogeography and conservation of Archangiopteris somai Hayata and A. itoi Hayata based on nucleotide variation of the atpB-rbcL intergenic spacer of chloroplast DNA. Fern Gaz., 16: 335–340.

Chiou, W.-L., Farrar, D. R., & Ranker, T. A. 2002. The mating systems of some epiphytic Polypodiaceae. Am. Fern J., 92: 65–79.

Chiou, W.-L., Huang, Y.-M., & Lee, P.-H. 2003. Mating systems of Cyatheaceae native to Taiwan. Pp. 485–489 in: S.Chandra & M.Srivastava (eds), Pteridology in the New Millennium. Netherlands: Springer, Netherlands.

Chiou, W.-L., Lee, P.-H., & Huang, Y.-M. 2003. The reproductive biology of gametophytes of homosporous ferns. Taiwan J. For. Sci., 18: 67–73.

Huang, Y.-M., Kuo, C.-Y., & Chiou, W.-L. 2003. Morphology of gametophytes and young sporophytes of Cibotium Kaulf. native to Taiwan. Taiwan J. For. Sci., 18: 163–170.

Huang, Y.-M., Won, S.-L., & Chiou, W.-L. 2003. The collection and storage of pteridophyte spores. Taiwan J. For. Sci., 18: 75–79.

Moore, S.-J., Hsieh, T.-H., Huang, Y.-M., & Chiou, W.-L. 2002. Diplazium maonense Ching, a poorly known species of the Athyriaceae (Pteridophyta) in Taiwan. Taiwan J. For. Sci., 17: 113–118.

Chiou, W.-L., Lin, J.-C., & Wang, J.-Y. 2001. Phenology of Cibotium taiwanense (Dicksoniaceae). Taiwan J. For. Sci., 16: 209–216.

Huang, Y.-M., Chiou, W.-L., & Lee, P.-H. 2001. Morphology of the gametophytes and young sporophytes of Cyatheaceae native to Taiwan. Taiwania, 46: 274–283.

Before 2000

Chiou, W.-L., Lee, P.-H., & Ying, S.-S. 2000. Reproductive Biology of Gametophytes of Cyathea podophylla (Hook.) Copel. Taiwan J. For. Sci., 15: 1–12.

Huang, Y.-M., Ying, S.-S., & Chiou, W.-L. 2000. Morphology of gametophytes and young sporophytes of Sphaeropteris lepifera. Am. Fern J., 90: 127–137.

Chiou, W.-L., Yang, K.-C., Yang, Y.-P., Hsu, K.-S., & Chen, S.-Y. 2000. Type specimens in the Herbarium of Taiwan Forestry Research Institute: Pteridophyta. I. Taiwan For. Res. Inst. Ext., 118: 1–92.

Chiou, W.-L., Farrar, D. R., & Ranker, T. A. 1998. Gametophyte morphology and reproductive biology in Elaphoglossum. Can. J. Bot., 76: 1967–1977.

Chiou, W.-L., & Farrar, D. R. 1997. Comparative gametophyte morphology of selected species of the family Polypodiaceae. Am. Fern J., 87: 77–86

Chiou, W. -L., & Farrar, D. R. 1997. Antheridiogen production and response in Polypodiaceae species. Am. J. Bot., 84: 633–640.

Chiou, W.-L. 1985. The gametophytes of Platycerium bifurcatum (Cav.) C. Yushania, 2: 19–23.

Chiou, W.-L., & Shieh, W.-C. 1984. The gametophytes of Adiantaceae in Taiwan. Yushania, 1: 1–10.